https://bitbucket.org/MakeHuman/makehum ... ing.py-179
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[None, None, None] #This data will be filled by hand in the exported file.
It's not documented anywhere else except for in the exporter code (but then again we did mention it was a development tool not ready for production yet). But it basically says that this property of skeletons has to be added in by hand, after the export.
Alternatively you should remove all the lines that say reference_plane from your skeleton file, but by doing that your bone rolls will be random, and probably not something useful.
This is no problem for loading poses, but it makes life more difficult when using the rig in tools like blender, as the bone axes will not have logical directions, which makes posing more difficult.